Prof Morgan Pratchett ~ Reef Research Leader: Coral Reef Ecology
ARC Centre of Excellence for Coral Reef Studies
- About
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- Teaching
- Interests
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- Research
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- Impacts of climate change on coral reef ecosystems
- Population outbreaks of crown-of-thorns starfish
- Population dynamics of scleractinian corals
- Ecological versatility in coral reef fishes
- Biology and ecology of butterflyfishes (Chaetodontidae)
Prof Morgan Pratchett has broad interests in population and community ecology of coral reef organisms, especially corals and fishes. His current research focuses on major disturbances that impact coral reef ecosystems, with a view to understanding differential responses and vulnerabilities among coral reef organisms.
Morgan has written many papers describing direct and indirect effects of coral bleaching and outbreaks of crown-of-thorns starfish, considering impacts on both coral assemblages and associated assemblages of coral reef fishes. He has also conducted extensive research on the biology and ecology of coral reef butterflyfishes.
- Honours
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- Fellowships
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- 2012 - Queensland Smart Futures Fellowship
- 2011 - ACU Titular Fellowship, University of Oxford
- 2008 - Sir Keith Murdoch Fellowship, American Australian Association
- 2008 - Sir John and Laurine Proud Research Fellowship, Lizard Island Reef Research Foundation
- 2008 - Churchill Fellowship, University of Newcastle, UK
- Memberships
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- Australian Coral Reef Society
- Fisheries Society of the British Isles
- Recording Secretary for International Coral Reef Society
- Publications
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These are the most recent publications associated with this author. To see a detailed profile of all publications stored at JCU, visit ResearchOnline@JCU. Hover over Altmetrics badges to see social impact.
- Journal Articles
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- Bollati E, D'Angelo C, Alerdice R, Pratchett M, Ziegler M and Wiedenmann J (2020) Optical feedback loop involving dinoflagellate symbiont and Scleractinian host drives colorful coral bleaching. Current Biology, 30 (13), pp. 2433-2445, DOI:10.1016/j.cub.2020.04.055.
- Burn D, Matthews S, Caballes C, Chandler J and Pratchett M (2020) Biogeographical variation in diurnal behaviour of Acanthaster planci versus Acanthaster cf. solaris. PLoS ONE, 15 (2), Article: e0228796, DOI:10.1371/journal.pone.0228796.
- Chase T, Pratchett M, McWilliam M, Hein M, Tebbett S and Hoogenboom M (2020) Damselfishes alleviate the impacts of sediments on host corals. Royal Society Open Science, 7, Article: 192074, DOI:10.1098/rsos.192074.
- Chase T, Pratchett M and Hoogenboom M (2020) Behavioral trade-offs and habitat associations of coral-dwelling damselfishes (family Pomacentridae). Marine Ecology Progress Series, 633, pp. 141-156, DOI:10.3354/meps13167.
- Cowan Z, Ling S, Caballes C, Dworjanyn S and Pratchett M (2020) Crown-of-thorns starfish larvae are vulnerable to predation even in the presence of alternative prey. Coral Reefs, 39, pp. 293-303, DOI:10.1007/s00338-019-01890-w.
- Graba-Landry A, Loffler Z, McClure E, Pratchett M and Hoey A (2020) Impaired growth and survival of tropical macroalgae (Sargassum spp.) at elevated temperatures. Coral Reefs, 339, pp. 475-486, DOI:10.1007/s00338-020-01909-7.
- McWilliam M, Pratchett M, Hoogenboom M and Hughes T (2020) Deficits in functional trait diversity following recovery on coral reefs. Proceedings of the Royal Society of London Series B, Biological Sciences, 287, Article: 20192628, DOI:10.1098/rspb.2019.2628.
- Nowicki J, Pratchett M, Walker S, Coker D and O'Connell L (2020) Gene expression correlates of social evolution in coral reef butterflyfishes. Proceedings of the Royal Society of London Series B, Biological Sciences, 287 (1929), DOI:10.1098/rspb.2020.0239.
- Pratchett M, Messmer V and Wilson S (2020) Size‑specific recolonization success by coral‑dwelling damselfishes moderates resilience to habitat loss. Scientific Reports, 10, Article: 17016, DOI:10.1038/s41598-020-73979-0.
- Pratchett M, McWilliam M and Riegl B (2020) Contrasting shifts in coral assemblages with increasing disturbances. Coral Reefs, 39, pp. 783-793, DOI:10.1007/s00338-020-01936-4.
- Pratchett M, Caballes C, Newman S, Wilson S, Messmer V and Pratchett D (2020) Bleaching susceptibility of aquarium corals collected across northern Australia. Coral Reefs, 39, pp. 663-673, DOI:10.1007/s00338-020-01939-1.
- Westcott D, Fletcher C, Kroon F, Babcock R, Plagányi E, Pratchett M and Bonin M (2020) Relative efficacy of three approaches to mitigate Crown‑of‑Thorns Starfish outbreaks on Australia’s Great Barrier Reef. Scientific Reports, 10, Article: 12594, DOI:10.1038/s41598-020-69466-1.
- More
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ResearchOnline@JCU stores 285+ research outputs authored by Prof Morgan Pratchett from 1999 onwards.
- Current Funding
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Current and recent Research Funding to JCU is shown by funding source and project.
Ian Potter Foundation - Lizard Island Reef Research Foundation - Crown-of-Thorns Starfish Research Grants
Effective sampling of Acanthaster cf. solaris to provide early-warning of new and renewed outbreaks
- Indicative Funding
- $62,727 over 2 years
- Summary
- This project aims to develop a holistic monitoring program that will not only help to deliver an early-warning syste, but will increase the viability of culling CoTS populations even during non-outbreak periods with the ultimate goal of reducing the frequency and severity of future outbreaks. Perhaps even more importantly, this project will deliver (for the first time) spatially and temporally explicit information about the structure and dynamics of crown-of-thorns starfish within the ?initiation zone? and in the lead-up to a new population irruption, which is a critical knowledge gap that limits understanding of the initiation and causes of population outbreaks.
- Investigators
- Morgan Pratchett, Andrew Hoey, Ciemon Caballes, Sven Uthicke and Jason Doyle in collaboration with Peter Doll and Janai Chandler (ARC Centre of Excellence for Coral Reef Studies and Australian Institute of Marine Science)
- Keywords
- Coral Reefs; Disturbance; Management; Monitoring
Australian Research Council - Centres of Excellence
ARC Centre of Excellence for Integrated Coral Reef Studies
- Indicative Funding
- $28,000,000 over 8 years
- Summary
- The overarching aim of the ARC Centre of Excellence for Integrated Coral Reef Studies is to provide the scientific knowledge necessary for sustaining ecosystem goods and services of the world's coral reefs, which support the livelihoods and food security of millions of people in the tropics. The Centre will enhance Australia's global leadership in coral reef science through three ambitious research programs addressing the future of coral reefs and their ability to adapt to change. A key outcome of the research will be providing tangible benefits to all Australians by bui8lding bridges between the natural and social sciences, strengthening capacity, and informing and supporting transformative changes in coral reef governance and management.
- Investigators
- Graeme Cumming, Ove Hoegh-Guldberg, Malcolm McCulloch, Peter Mumby, Sean Connolly, John Pandolfi, Bob Pressey, Andrew Baird, David Bellwood, Joshua Cinner, Sophie Dove, Maja Adamska, Mia Hoogenboom, Geoff Jones, Mike Kingsford, Ryan Lowe, Mark McCormick, David Miller, Philip Munday, Morgan Pratchett, Garry Russ and Tiffany Morrison in collaboration with Janice Lough, David Wachenfeld, Stephen Palumbi, Serge Planes and Philippa Cohen (ARC Centre of Excellence for Coral Reef Studies, The University of Queensland, The University of Western Australia, College of Science & Engineering, Australian National University, College of Public Health, Medical & Vet Sciences, Australian Institute of Marine Science, Great Barrier Reef Marine Park Authority, Stanford University, Centre National de la Recherche Scientifique and WorldFish)
- Keywords
- coral reef ecosystems; Climate Change Adaptation; ecological resilience; biodiversity goods and services; social-ecological dynamics
Department of the Environment and Energy - Director of National Parks - Tender
Coral Sea Marine Park Coral Reef Health Survey (2021)
- Indicative Funding
- $354,402 over 1 year
- Summary
- The Coral Sea Marine Park (CSMP) covers approximately 990,000 km2 and includes over 30 individual reef systems that are surrounded by deep waters. In March 2020 extensive bleaching of corals was recorded across shallow reef habitats within the CSMP. This project will use surveys of benthic, fish and invertebrate communities to assess the impacts of the 2020 bleaching event on coral reefs within the CSMP.
- Investigators
- Andrew Hoey, Morgan Pratchett and Hugo Harrison (ARC Centre of Excellence for Coral Reef Studies)
- Keywords
- Coral Reef; Coral Bleaching; Marine Mark Management; Reef Fish; Coral
Department of the Environment and Energy - Director of National Parks - Contract Research
Coral Sea - Coral Reef Health Multi Year Project
- Indicative Funding
- $45,455 over 2 years
- Summary
- The principal output of this project will be providing spatial and temporal analyses of hydrodynamic patterns in the Coral Sea Marine Park (CSMP) to facilitate planning and establishment and/or revisions of existing management plans. This project will inform patterns of connectivity and circulation flow across the broader CSMP, and between CSMP and Great Barrier Reef Marine Park using existing modelling platforms.
- Investigators
- Morgan Pratchett, Hugo Harrison and Severine Choukroun (ARC Centre of Excellence for Coral Reef Studies)
- Keywords
- Coral Reefs; Hydrodynamic Modeling; Coral Sea Marine Park; Connectivity
Department of the Environment and Energy - National Environmental Science Program (NESP) - Tropical Water Quality Hub (TWQ Hub)
Innovations in COTS Control on the Great Barrier Reef
- Indicative Funding
- $2,900 over 1 year (administered by Reef & Rainforest Research Centre)
- Summary
- Innovations in COTS Control on the Great Barrier Reef. For decades, crown-of-thorns starfish (COTS) research effort has been fragmented and failed to inform improvements in the effectiveness of COTS control. However, under the NESP Tropical Water Quality Hub?s Integrated Pest Management (IPM) research program, a more collaborative and targeted approach has harnessed research effort and enabled delivery of significant improvements in effectiveness and efficiency of COTS control, while also looking to develop methods for the future. Wholescale adoption of this IPM approach by the Great Barrier Reef Marine Park Authority (GBRMPA) and operators is already demonstrably saving live coral cover on the Great Barrier Reef (GBR).
- Investigators
- Suzanne Long, Frederieke Kroon, Sven Uthicke, Jason Doyle, Russ C Babcock, Morgan Pratchett and Peter Mumby in collaboration with David Logan, Paulina Kaniewska, David Westcott, Cameron Fletcher, Eva Plaganyi and Scott Condie (Reef and Rainforest Research Centre, Australian Institute of Marine Science, Commonwealth Scientific & Industrial Research Organisation, ARC Centre of Excellence for Coral Reef Studies, The University of Queensland and Parks Australia)
- Keywords
- Crown-of-thorns starfish; Integrated Pest Management; Great Barrier Reef
Australian Museum - Crown-of-Thorns Starfish research grants
Early detection of Crown-of-Thorns starfish recruitment using settlement traps and genetic analyses
- Indicative Funding
- $22,727 over 1 year
- Summary
- Major knowledge gaps remain around the settlement and early post-settlement stages of Crown-of-Thorns starfish (CoTS). Early detection of large settlement pulses provides our best chance to potentially stop future CoTS outbreaks. Our recent work has shown that CoTS larvae settled to our custom-built settlement traps. In order to detect settlement, genetic analyses were developed to analyse the samples. The aim is to additionally use new innovative analyses that will allow us to compare settlement rates at different time points, reefs, sites and depths. More traps will also be deployed during the next spawning season to improve our understanding of settlement preferences and test the traps as an early warning system on reefs not subject to outbreaks yet.
- Investigators
- Vanessa Messmer and Morgan Pratchett in collaboration with Jason Doyle and Sven Uthicke (ARC Centre of Excellence for Coral Reef Studies and Australian Institute of Marine Science)
- Keywords
- Crown-of-Thorns Starfish; Genetic Analysis; Early detection; digital droplet PCR; settlement traps; CoTS ourbreaks
Department of the Environment and Energy - National Environmental Science Program (NESP) - Tropical Water Quality Hub (TWQ Hub)
Crown-of-thorns starfish: surveillance and life history
- Indicative Funding
- $150,000 over 3 years (administered by CSIRO)
- Summary
- This project supports the implementation of the NESP Crown-of-Thorns Starfish Integrated Pest Management Project's management and research strategy by i) describing ecological parameters key to parameterizing the tools underpinning the management strategy, ii) analysing existing data sets to assess management performance and describe the interaction between CoTS and bleaching, and , iii) scoping new technologies and surveillance strategies for detecting and monitoring primary and secondary outbreaks. The work will involve a combination of field ecological research, statistical analysis of existing data, technology and strategy development (the latter through workshops) and inform decision making through incorporation into decision support tools and recommendations.
- Investigators
- David Westcott and Morgan Pratchett in collaboration with Cameron Fletcher, Russ C Babcock, Frederieke Kroon, Sven Uthicke, Jason Doyle, Charlotte Johansson, Vanessa Messmer, Peter Mumby and Pascal Craw (Commonwealth Scientific & Industrial Research Organisation, Australian Institute of Marine Science, ARC Centre of Excellence for Coral Reef Studies and The University of Queensland)
- Keywords
- Coral Reefs; Management; Disturbance; Surveillance; Biology; Acanthaster
Department of the Environment and Energy - Director of National Parks - Tender
Coral Reef Health in the Coral Sea Marine Park
- Indicative Funding
- $968,346 over 2 years
- Summary
- A number of coral reefs occur in the Australian Marine Parks estate, particularly within the Coral Sea Marine Park (CSMP). In 2016 and 2017, coral reefs in the Great Barrier Reef Marine Park and CSMP were impacted by a mass global coral bleaching event. Regular surveys of coral bleaching and associated reef health in the CSMP are needed to inform future marine park management over the next three years to either enable comparisons against historical baselines where bleaching has previously occurred, or set new baselines for areas that aren?t bleached but may be in the future.
- Investigators
- Morgan Pratchett, Andrew Hoey, Hugo Harrison, Andrew Baird, Tom Bridge, Daniela Ceccarelli and Jean-Paul Hobbs (ARC Centre of Excellence for Coral Reef Studies and Curtin University of Technology)
- Keywords
- Coral Reefs; Monitoring; Coral Bleaching; Coral Sea Marine Park
Department of the Environment and Energy - National Environmental Science Program (NESP) - Tropical Water Quality Hub (TWQ Hub)
Implementation of the Crown of Thorn Starfish research strategy: regional strategies
- Indicative Funding
- $85,000 over 3 years (administered by CSIRO)
- Summary
- Crown-of-Thorns starfish (COTS) management requires decisions be made about where to invest control effort in order to achieve regional scale objectives, e.g. to modify COTRS spread or to facilitate coral recovery. However, no objective basis for choosing sites exists, beyond their economic importance, meaning that achieving these important goals is largely left to chance. This project will leverage past and current research to develop a decision support framework to assist in prioritizing reefs for control and for assessing alternative strategies by integrating the full range of ecological and management information available. Field and desktop research will also be conducted to fill critical knowledge gaps and to assess the prospects of new control technologies.
- Investigators
- David Westcott and Morgan Pratchett in collaboration with Peter Mumby, Cameron Fletcher, Russ C Babcock, Karlo Hock, Scott Condie, Rebecca Gorton, Eva Plaganyi-Lloyd, Lone Hoj, Maria Byrne, Frederieke Kroon, P Costello, Sven Uthicke, Jason Doyle, Frances Patel, Robin Beaman and Matthew Dunbabin (Commonwealth Scientific & Industrial Research Organisation, The University of Queensland, ARC Centre of Excellence for Coral Reef Studies, Australian Institute of Marine Science, The University of Sydney, College of Science & Engineering and Queensland University of Technology)
- Keywords
- Acanthaster; Great Barrier Reef; Integrated Pest Management; Tagging; Movement; Field surveys
Fisheries Research & Development Corporation - Annual Competitive Round
Establishing baselines and assessing vulnerability of commercially harvested corals across northern Australia
- Indicative Funding
- $384,300 over 5 years (administered by JCU)
- Summary
- This project is critically important to provide necessary data on the distribution, abundance and biology of commonly harvested corals across northern Australia, both to ensure that current harvest levels and practices are sustainable and address legislative requirements for harvesting and exporting species listed under the Convention on International Trade in Endangered Species (CITES).
- Investigators
- Morgan Pratchett and Vanessa Messmer in collaboration with Stephen Newman (ARC Centre of Excellence for Coral Reef Studies and WA Department of Fisheries)
- Keywords
- Corals; aquarium fishery; Stock Assessment; Population Structure; Vulnerability
Australian Museum - Crown-of-Thorns Starfish research grants
Distribution, abundance and reproductive behaviour of non-outbreak populations of Acanthaster cf. solaris
- Indicative Funding
- $23,004 over 1 year
- Summary
- This project will investigate the relative importance of diet regime and reproductive effort on the overall health of the Crown-of-Thorns Starfish. The aim is to determine the underlying causes and mechanisms that lead to population crashes among COTS at the end of major outbreaks, filling the knowledge gap on what naturally regulates abundance of COTS. This will provide valuable data that will be of great help to the selection of areas to be controlled, thereby increasing the efficiency and effectiveness of control activities, ultimately assisting in the management of this species and preventing coral loss.
- Investigators
- Morgan Pratchett and Andrew Hoey (ARC Centre of Excellence for Coral Reef Studies)
- Keywords
- Acanthaster cf. solaris (Acanthasteridae; Management; distu7rbance; Surveillance; Biology
Department of Parks and Wildlife - Western Australia - Contract Research
Abundance, Species Identify and Resilience of Inshore Corals in the Dampier Archipelago
- Indicative Funding
- $150,000 over 3 years
- Summary
- The study will undertake fisheries-independent assessments of the abundance and composition of corals in near shore environments across the Dampier Archipelago. Importantly, detailed sampling will be combined with genetic sampling and analysis to unequivocally establish the species identity of corals, as required for aligning catch data with scientific information on the vulnerability of different coral species. This project will provide important baseline information on the ecological and economic importance of near shore coral assemblages and contribute to the development of monitoring protocols necessary to ensure on-going harvesting is sustainable, even in the face of climate change.
- Investigators
- Morgan Pratchett in collaboration with Vanessa Messmer and Cassandra Thompson (ARC Centre of Excellence for Coral Reef Studies)
- Keywords
- Corals; Fisheries; Population Deography; Marine Aquarium Industry; Stock Assessment
- Supervision
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Advisory Accreditation: I can be on your Advisory Panel as a Primary or Secondary Advisor.
These Higher Degree Research projects are either current or by students who have completed their studies within the past 5 years at JCU. Linked titles show theses available within ResearchOnline@JCU.
- Current
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- Identifying key indicators to quantify ecological recovery and adaptation of coral reefs across space and time (PhD , Secondary Advisor/AM)
- Interactions between fish and coral reefs: the influence of habitat degradation on fish communities (PhD , Secondary Advisor/AM)
- Phylogenomics and Whole Genome Assembly of Marine Angelfishes: The Evolutionary History of an Iconic Reef Fish Family (PhD , Secondary Advisor/AM)
- Development and Application of Novel Genetic Tools for the Investigation of Genetic Diversity and Age Structure of Crown-of-Thorns Seastar on the Great Barrier Reef (PhD , Secondary Advisor/AM)
- Changes in behaviour and sociality of Chaetodon butterflyfish linked to population declines and coral loss (Masters , Primary Advisor)
- Effective Sampling of Acanthaster cf.Solaris to provide early Warning of New and Renewed Outbreaks (Masters , Primary Advisor/AM/Adv)
- The effects of macroalgal stands on coral demographics on the GBR (PhD , Secondary Advisor)
- Vulnerability and Resilience of Coral Assemblages in Australia's CoralSea Marine Park (PhD , Primary Advisor/AM/Adv)
- Biology and Ecology of Australian Aquarium Corals: Information needed to Secure the Sustainability of ongoing Wild Harvest of Scleractinian Corals (PhD , Primary Advisor/AM/Adv)
- The Effect of Ocean Warming on the Behaviour and Fitness of Post-settlement Crown-of-Thorns Starfish (Acanthaster cf. solaris) (PhD , Primary Advisor/AM/Adv)
- Habitat suitability and patterns of larval settlement for crown-of-thorns starfish (Acanthaster cf. solaris) in the western Pacific (PhD , Primary Advisor/AM/Adv)
- Dynamics of Reef-building corals along the north-western Australian coastline (PhD , Secondary Advisor)
- The Role of Hybridisation in the Evolution of Coral Reef Fishes (PhD , Advisor Mentor)
- Completed
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- Predation on the early life stages of the crown-of-thorns starfish (Acanthaster cf. solaris) (2017, PhD , Primary Advisor)
- Neural and ecological basis of pair bonding in butterflyfishes (F: Chaetodontidae) (2017, PhD , Primary Advisor)
- Effects of habitat versus fisheries management on spatio-temperal variation in fish assemblages on the Great Barrier Reef (2017, PhD , Primary Advisor)
- Effects of climate change-induced thermal stress and habitat degradation to the biodiversity and species composition of coral-associated invertebrates (2016, PhD , Secondary Advisor)
- Environmental influences on the reproductive biology and early life history of the crown-of-thorns starfish (2017, PhD , Primary Advisor)
- Adaptive strategies in reef-building corals (2017, PhD , Secondary Advisor)
- Vectors and environmental drivers of coral disease dynamics on the Great Barrier Reef (2017, PhD , Secondary Advisor)
- Causes and consequences of natural hybridisation among coral reef butterflyfishes (Chaetodon: Chaetodontidae) (2018, PhD , Secondary Advisor)
- Variation in structure and function of reef fish assemblages among distinct coral habitats (2018, PhD , Secondary Advisor)
- Modelling tools for the management of crown-of-thorns starfish (Acanthaster cf. solaris) outbreaks on Australia’s Great Barrier Reef (2019, PhD , Primary Advisor/AM/Adv)
- Spatial and temporal variation in movement patterns and behaviour of a large, coral reef mesopredator, Plectropomus leopardus, on the Great Barrier Reef, Australia (2019, PhD , Primary Advisor/AM/Adv)
- The persistence of Sargassum communities on coral reefs: resilience and herbivory (2019, PhD , Secondary Advisor)
- Effects of coral-dwelling damselfishes' abundances and diversity on host coral dynamics (2019, PhD , Secondary Advisor)
- Spatial and temporal variation in the demography of early-stage juvenile Pacific crown-of-thorns starfish (Acanthaster cf. solaris) from Australia’s Great Barrier Reef (2020, PhD , Primary Advisor/AM/Adv)
- Temporal and Spatial Variation in the Growth of Branching Corals (2016, PhD , Primary Advisor)
- The effect of increasing temperature on algae-fish interactions on coral reefs (2020, PhD , Secondary Advisor)
- Differential susceptibility of corals to major versus routine disturbances in Australia’s Coral Sea (2021, Masters , Primary Advisor/AM/Adv)
- Temperature preferences of tropical fishes and the influence of local abiotic and biotic factors (2020, PhD , Secondary Advisor)
- Data
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These are the most recent metadata records associated with this researcher. To see a detailed description of all dataset records, visit Research Data Australia.
- Pratchett, M. (2020) Gamete concentrations and gonad index for Pacific crown-of-thorns starfish. James Cook University
- Graba-Landry, A. (2020) Graba-Landry PhD thesis: The effect of increasing temperature to algae-fish interactions on coral reefs. James Cook University
- Burn, D. (2020) Biogeographical variation in diurnal behaviour of Acanthaster planci versus Acanthaster cf. solaris. James Cook University
- Hughes, T. (2019) Global warming impairs stock-recruitment dynamics of corals. James Cook University
- Richardson, L. (2018) PhD thesis data: Variation in structure and function of reef fish assemblages among distinct coral habitats, by Laura E. Richardson (2018). James Cook University
- Hughes, T. (2018) Global warming transforms coral reef assemblages. James Cook University
- Blowes, S. (2017) Data from: Heterospecific aggression and dominance in a guild of coral-feeding fishes: the roles of dietary ecology and phylogeny. James Cook University
- Lawton, R. (2017) Data from: High gene flow across large geographic scales reduces extinction risk for a highly specialised coral feeding butterflyfish. James Cook University
- Pratchett, M. (2017) Captive movement rates of the Pacific crown-of-thorns starfish, Acanthaster solaris. James Cook University
- Cowan, Z. (2017) Predation on settling crown-of-thorns starfish larvae by benthic predators. James Cook University
- Collaboration
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The map shows research collaborations by institution from the past 7 years.
Note: Map points are indicative of the countries or states that institutions are associated with.- 5+ collaborations
- 4 collaborations
- 3 collaborations
- 2 collaborations
- 1 collaboration
- Indicates the Tropics (Torrid Zone)
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